Coral reefs are one of the most biodiverse ecosystems on Earth and have inspired the development of some of the most far-reaching theories in ecology. These ecosystems are distributed throughout the tropics and often dominate the shallow seas. They are also important for many millions of people worldwide yet are under increasing threat from climate change and more direct anthropogenic disturbance. This module aims to provide a solid grounding in coral reef biology, ecology and evolution, with a focus on corals and reef fishes, building on broad ecological principles laid down in previous years. Students will apply this understanding to evaluate threats and their potential solutions, developing an appreciation of the precarious nature of the most complex habitat in the oceans. Specifically, the students will explore how and where coral reefs have emerged through time and adapted to life in the oceans, the delicate balance of interactions that allow their enormous variety of species to coexist, and emerging threats and solutions to their continued existence.